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Summer course in embryology of marine invertebrates - 2, 2016

The White Sea Biological station (WSBS) of the M.V. Lomonosov Moscow State University (MSU) offers 3-week field course in embryology of marine invertebrates 13 June – 03 July 2016.



The White Sea Biological Station is located on the wild and scenic coast of Kandalaksha Bay of the White Sea (66° 34' N, 33° 08' E) exactly on the Polar Circle. It is a remote and isolated academic settlement accessible only by boat from the nearest village Poyakonda. Station facilities comprise a number of recently upgraded laboratories with recirculating seawater system, cLSM, and SEM. The station has good diving facilities and a fleet of motorboats and small vessels designed for benthic and pelagic sampling.

See the detailed information about the station:


Tentative list of instructors

Carmen Andrikou (Sars International Center for Marine Molecular Biology, NORWAY)
Lev Beloussov (Dept. of Embryology, MSU, RUSSIA)
Nataliya Budaeva (University Museum of Bergen, NORWAY)
Alexander Ereskovsky (Station marine d'Endoume, Aix-Marseille Université, FRANCE; Dept. of Embryology, Saint-Petersburg State University, RUSSIA)
Grigory Genikhovich (Dept. of Molecular Evolution and Development, University of Vienna, AUSTRIA)
Andreas Hejnol (Sars International Center for Marine Molecular Biology, NORWAY)
Yulia Khramova (Dept. of Embryology, MSU, RUSSIA)
Igor Kosevich (Lab. of Developmental Biology, MSU, RUSSIA)
Olga Kotenko (Dept. of Invertebrate Zoology, St. Petersburg State University, RUSSIA)
Yulia Kraus (Dept. of Evolutionary Biology, MSU, RUSSIA)
Stanislav Kremnyov (Dept. Of Embryology, MSU, RUSSIA)
Andrey Lavrov (Dept. of Invertebrate Zoology, MSU, RUSSIA)
Vladimir Malakhov (Dept. of Invertebrate Zoology, MSU, RUSSIA)
Mark Martindale (The Whitney Laboratory for Marine Bioscience&Seahorse Key Marine Laboratory, University of Florida, USA)
Denis Nikishin (Group of Embryophysiology; Koltzov Institute of Developmental Biology, RUSSIA)
Andrey Ostrovsky (Dept. of Paleontology, University of Vienna, AUSTRIA & Department of Invertebrate Zoology, St Petersburg State University, RUSSIA)
Andrey Prudkovsky (Dept. of Invertebrate Zoology, MSU, RUSSIA)
Nadezhda Rimskaya-Korsakova (Dept. of Invertebrate Zoology, MSU, RUSSIA)
Andreas Wanninger (Dept. of Integrative Zoology, University of Vienna, AUSTRIA)
Maria Semenova (Dept. of Embryology, MSU, RUSSIA)
Alexander Tzetlin (WSBS, MSU, RUSSIA)

Course description

The Course will be lab-oriented with daily lectures on various aspects of invertebrate development. The students will learn the methods of culturing of marine invertebrates and will be taught how to document the development using traditional illustration techniques and digital photography and video. The students will also carry out simple individual projects on development of a particular species. The main goals of the individual projects are to demonstrate various methods and approaches in experimental embryology such as immunocytochemistry and confocal microscopy and pharmacological modulation.

We plan to study the development of approximate 35 species from 10 invertebrate phyla (Porifera; Cnidaria; Annelida; Mollusca; Lophophorata; Nemertea; Arthropoda; Nematoda; Echinodermata; Tunicata). All species are easy accessible in close vicinity of the station and can be collected either in the intertidal one or by the scuba divers. Similar course was organized at WSBS in 2010, please find more information about it here.

Link for Presentation of the Course

You can find the tentative programme of the Course here:

Скачать   EMBRYO_PRO_WSBS2016.pdf
      768.68 KB




Applications are invited preferably from graduate students however well qualified undergraduate student as well as early career post-docs and researchers interested in embryology of marine invertebrates, may also apply.

LANGUAGE OF INSTRUCTION: ENGLISH. No formal test results are required (such as TOEFL), however, all applicants are expected to have good English language skills.

Student costs

TUITION: The course tuition is 400 Euro, however, it may be partially covered by the WSBS, for students facing financial constraints.

WSBS is an old academic settlement with more than 70 years of history. Student's dorm is a wooden house with wood-burning stoves and outdoor sanitary facilities and toilets. Students enrolled in the class should expect simple living conditions, which however are well compensated by beautiful scenery and untouched nature.
WSBS provides dormitory accommodation for students enrolling in classes (rooms for 4–6 people) for free, however small compensation fee covering cleaning prior to departure and bed linen (one-time payment; approximately 15 Euro) is applicable to all WSBS visitors.

The full-board meal plan is obligatory for all persons living in dormitories. Full board consists of three meals a day and costs about 8 Euro per day. Vegetarian options are available, however we can not provide Kosher or Halal meals, due to isolated location of the field station.

All students enrolled in the class are obligated to have personal medical insurance including medical evacuation coverage.

Train (Moscow-Kandalaksha), second class round trip fare: approximately 190 Euro.
Bus (Kandalaksha-Poyakonda), round trip: approximately 40 Euro.
Boat from Poyakonda to WSBS will be arranged for all participants: approximately 7 Euro per trip.
All payments must be done at the field station in cash in Russian Roubles (RUR). We do not accept credit or debit cards.

International students are required to have entry visas to travel to the Russian Federation. Russian entry visas can be obtained at a local Russian Embassy or Consulate. All supporting materials will be sent upon acceptance to the course.

Students facing financial difficulties may apply for financial support from WSBS. Financial aid may include full or partial waiver of tuition. We do not provide coverage for the transportation coasts.

Application process

1) Cover page with personal data (please download from this web page, complete, sign, scan and return as TIF, JPEG or PDF file )


2) A one-page personal statement, describing how this course will help advance applicant's educational goals and career.
3) Curriculum Vitae (please, do not include reprints).
4) One letter of support from a person most familiar with applicant's work. Letter of support may be sent directly from the referee's e-mail address. Alternatively, a signed letter may be scanned and sent electronically by the applicant.
5) Statement of financial need, if applying for support from the WSBS.

Complete applications should be sent electronically to the following e-mail address: with the following subject: “EmbryoCourse+your last name”.

Maximum 10 students.

Application deadline: February 15, 2016
Acceptance announcement: March 15, 2016

Applications will be acknowledged once they have been processed.
For additional information contact

Web-site of the course: 

Dear colleagues,

Could you please distribute the following information to potential applicants?
Thank you in advance!

Скачать announcement_short_eng_2016.docx  18.74 KB




The school is supported by: 

  • Report
    Report on the White Sea Biological Station of Moscow State University (WSBS MSU) held the Second International Summer Course on the embryology of marine invertebrates in June 2016 (12/06-03/07)


Photos: 59


White Sea Biological Station, Poselok Primorskiy,
Republic Karelia, RUSSIA

Tel/fax: +7 (815) 33-64-516

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