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Dr. Grigory Kolbasov

Leading Scientist
Morphology, phylogeny, biology and evolution of crustaceans, study and monitoring of marine benthic communities.

leading scientist

Business address:                                                             
Moscow State University                                    
Biological Faculty,                                              
White Sea Biological Station,                             
Moscow 119992, RUSSIA
+7(495)9394233 (tel.)
+7(495)9394233 (fax)

Biographical data:
Present Rank: Doctor of Science in Zoology, leading researcher
Date of Birth: July 17, 1967
Place of Birth: Moscow, Russia

Secondary School education:
from September 1974 to June 1984 (Moscow secondary schools No.398, 57).

Higher education:
• From September 1984 to July 1989 – studies at Moscow State University (department of biology). Master of Science. Zoology.
• Diploma "with honors". The topic of diploma thesis: “Sponge-inhabiting barnacles of the genus Acasta, systematics, morphology, ecology and distribution”. Supervisor: Dr G.B. Zevina (Moscow State University, Moscow, Russia).

Post-graduate courses, defended degree theses:
• From October 1989 to October 1992 – a post-graduate student at Moscow State University.
• In October 1992 – defended a candidate dissertation. The topic of Ph.D. thesis: “Sponge-inhabiting barnacles, systematics, morphology, evolution, ecology and distribution”. Supervisor: Dr. G.B. Zevina.
• In October 2002 – defended Doctorate dissertation. The topic of Doctorate thesis: “Burrowing barnacles of the superorder Acrothoracica, systematics, phylogeny, morphology, ecology and distribution”.

Professional employment record:
• from 2006 to present – leading researcher of the White Sea Biological Station of Biological Faculty of Moscow State University (Moscow, Russia);
• 1999-2006 – senior researcher of the White Sea Biological Station of Biological Faculty of Moscow State University (Moscow, Russia);
• 1998-2003 Deputy Chair of the White Sea Biological Station
• 1992-1999 – researcher of the White Sea Biological Station of Biological Faculty of Moscow State University (Moscow, Russia).

Subjects of scientific work:
• taxonomy, systematics, phylogeny, morphology, and ecology of marine crustaceans;
• investigation of the biology, ecology, distribution (monitoring) of benthic communities of Arctic Seas.

Field of scientific interests:
• benthic ecology
• biodiversity and biomonitoring
• biogeography
• cladistics and phylogeny
• coral reef ecology
• deep-sea ecology
• hydrobiology
• marine zoology
• parasitology of marine organisms
• taxonomy and systematics
• zoogeography

Professional skills:
• experience in taxonomical, morphological and phylogenetic works;
• experience in ecological studies of meiobenthos;
• methods of statistical treatment and database creating;
• method of sampling and treatment of meiobenthic samples;
• methods of sampling and treatment of macrobenthic and meiobenthic samples during vessel cruises and diving surveys;
• field experience in research vessel and diving surveys;
• methods of scanning and transmission electron microscopy;
• professional SCUBA diver (possessing a Russian professional diving license)

Honors and awards:
• 1989 – Diploma of Moscow State University “with honor”;
• 1989-1992 – Soros’ Ph.D. student and researcher (grant of the International Soros’ Science Education Program);
• 2007 – awarded Shuvalov Prize of Moscow State University with medal for best Doctorate thesis.

Teaching experience (1989 – present):
Practical course on Invertebrate Zoology for first year students of Moscow State University of Biological Faculty
Practical courses on Invertebrate Zoology at the White Sea Biological Station for students of Moscow State University.
Lection course on phylogeny and morphology of crustaceans for fourth year students of Department of Invertebrate Zoology of Biological Faculty of Moscow State University.
Lection course on the bioinvasions for third year students of Department of Biogeography of Geographical Faculty of Moscow State University
Lection course on arthropodology for fourth year students of Department of Invertebrate Zoology of Biological Faculty of Moscow State University.

Ph.D. students and graduate students:
Alexandra S. Petrunina, dept. of Invertebrate Zoology, Ph.D. thesis “Microparasitic crustaceans of class Tantulocarida. Phyllogeny, systematics, morphology, development and life circle” (2008-2012). Thesis was defended in 2012.
• 2012-2014 - graduate student of Department of Invertebrate Zoology of MSU (Magister degree), Sokolova A.M., theme "Macrofauna associated with sponge Niphates olemda Laubenfels, 1936 (Demospongiae: Haplosclerida) from Nhatrang Bay, South Vietnam".

Referee for: Crustaceana, Jour. Crustacean Biol.; J Mar. Biol. Ass. U.K.; Molecular Phylogenetics and Evolution; Raffles J. Zool.; Zoologischer Anzeiger; Zootaxa and other journals.

Participation in scientific programs and expeditions:
1. 1987, 1988 – expeditions in Far East of Soviet Union (Kurile Islands, Japan Sea), studies of benthic communities and cirripedes.
2. 1989 – expedition in South Vietnam, studies of marine fouling and cirripedes
3. 1996, 1998, 2000, 2007 – guest scientist at Copenhagen University, Denmark, Zoological Institute, studies on morphology, phylogeny and systematics of Crustacea.
4. 2011, 2012, 2013, 2014, 2015, 2016, 2017, 2018, 2020 - scientific trips in different sites of Taiwan (binary grants), studies of symbiotic crustaceans of tropical Pacific

Personal grants (Principal investigator PI):
1. 1997-1998. Russian Foundation for Basic Research: “System, morphology, evolution and biology of the cirripedes of the order Acrothoracica and their role in the superorder Cirripedia origination”.
2. 1999-2001. Russian Foundation for Basic Research: “Morphology and biology of crustaceans of the subclass Facetotecta, their systematics and relationships with other taxa of the class Thecostraca”.
3. 1999. Lerner Gray Fund, American Museum of Natural History, travel grant on study of crustaceans of the subclass Facetotecta.
4. 2002-2004. Russian Foundation for Basic Research: “Crustaceans of the subclass Ascothoracida: the morphology and systematic, relationships with other taxa of the class Thecostraca”.
5. 2006-2008. Russian Foundation for Basic Research: “The parasitic microcrustaceans of the class Tantulocarida. Morphology, systematic, development and ecology”.
6. 2009-2011. Russian Foundation for Basic Research: “Cypris larva – a clue to understanding of cirripede evolution (Crustacea: Cirripedia). Morphology, anatomy, phylogeny”.
7. 2008. Russian Foundation for Basic Research: Publishing of a book “Acrothoracica – the burrowing barnacles”.
7. 2009. Russian Foundation for Basic Research, participant in conference of 'Integrative Biology Society', symposium 'Biology of parasitic crustaceans'.
8. 2011. Russian Foundation for Basic Research, expedition for collecting of larvae of crustaceans of classes Thecostraca and Tantulocarida in Northern Seas of Russia.
9. 2011-2013. Russian Foundation for Basic Research and MOST, binary Russian-Taiwanese grant 11-04-92002-NNS_а, 'Reconstruction of the phylogeny of barnacles of the superorder Acrothoracica (Crustacea, Cirripedia) with using morphological and molecular approaches. New synthetic systematics of the Acrothoracica.
10. 2014-2016. Russian Foundation for Basic Research and MOST, binary Russian-Taiwanese grant 14-04-92002 NNS_а, Barnacles (Cirripedia, Balanidae, Acastinae) - symbionts of sponges and cnidarians. Systematics, molecular phylogeny and evolution of symbiontic relations.
11. 2015-2017. Russian Foundation for Basic Research, 15-04-00259а, "Phylogeny of the crustaceans of the classes Thecostraca and Tantulocarida".
12.2017-2020 Russian Foundation for Basic Research RFBR, 17-54-52006 MNT_a, Ascothoracida (Thecostraca, Crustacea) - parasites of cnidarians and echinoderms. Recontruction of the phylogeny with using morphological and molecular approaches and evolution of the parasitism within this group.
13. 2018-2020 Russian Foundation for Basic Research, 18-04-00624а, A hundred years without victory or Y-larvae - an old enigma in marine biology. 2018-2020
14. 2021-2023. Russian Foundation for Basic Research RFBR, 21-54-52003 MNT_a, Parasitic Thecostraca in the coral reef communities: diversity, life history, relationships of major taxa, host-parasite relations

Shared external grants (Co-investigator):
1. 1995-1997. Russian Foundation for Basic Research: “Study of space-temporal structure and trophic nets in coastal marine ecosystems. Factors integrating marine ecosystems”.
2. 1998-2000. Russian Foundation for Basic Research: “Dynamics of the seashore biotops and the cyclic of the successions. The choice of the parameters for ecological monitoring”.
3. 2009-2012. Russian Federal Agency of Science and Innovations (02.740.11.0280, 02.740.11.0875 and P1291) and Program of Leading Scientific Schools (NSh-4456.2010.4)

Participation in conferences/symposiums:
1. 1995. Conference "Problems of study, rational exploration and preservation of natural resources of the White Sea" (Kandalaksha, 1995).
2. 1996 – 2006. Annual Conference on Marine Biology at the White Sea Biological Station (White Sea Biological Station, Murmansk district).
3. 1998. The Fourth International Crustacean Congress (Amsterdam, the Netherlands, 1998).
4. 2007. Second International conference "Ecological studies of the White Sea organisms" (Kartesh, Biological Station of Zoological Institute, Karelia)
5. The commemorative readings in honor of eminent zoologist and parasitologist Prof. V.L.Vagin (1907-1984) „Marine zoology and parasitology: investigations and discoveries” (Kazan, Russia, 2007).
6. 2008. Eight International Symposium on Larval Biology (Lisbon, Portugal, 2008).
7. 2008. International Symposium “Advances in Crustacean Phylogenetics” (Rostock, Germany, 2008).
8. 2009. Annual Meeting of Society for Integrative and Comparative Biology (Boston, USA, 2009).
9. 2010. 14th International Meiofauna Conference, Ghent, Belgium.
10. 2011. Russian conference on echinoderms. Institute of Oceanology RAS, Moscow.
11. 2011. 11th International Conference on Copepoda, Merida, Mexico.
12. 2011. World Conference on Marine Biodiversity, Aberdeen, UK.
13. 2012. Crustacean Society Summer Meeting and the 10th Colloquium Crustacea Decapoda Mediterranean, Athens, Greece.
14. 2013. Scientific conference "Marine biology, geology, oceanology - interdiciplinary studies on marine stations". White Sea Biological Station of MSU
15. 2013. Summer Meeting of the Crustacean Society, San José, Costa Rica.
16. 2013. Scientific conference "Molecular genetics approaches in taxonomy and ecology" Rostov-na-Donu
17. 8th International Crustacean Congress, 18-24 August 2014, Frankfurt, Germany, 102.
18. The 3rd International Congress on Invertebrate Morphology (ICIM-3). Berlin. August 2014. Germany. , Berlin, Germany
19. 19-23 July 2015 – The Crustacean Society mid-year meeting, with the International Astacology Assoc., Australian Museum, Sydney, New South Wales, Australia, 46.
20. 10-13 August 2017 -  XIth INTERNATIONAL LARVAL BIOLOGY SYMPOSIUM , Honolulu, Hawaii, USA
21. 18-23 August 2017. 4th International Congress on Invertebrate Morphology (ICIM4)/IV, Moscow, Russia
22. May 22-25, 2018. 9th International Crustacean Congress, Washington, USA, Washington, DC
23. 6 November 2018. 20th Anniversary of RFBR-MOST Partnership Conference, Taipei, Taiwan
24. NOVEMBER 19-21, 2018. MARINE BIOLOGY, GEOLOGY, OCEANOLOGY: INTERDISCIPLINARY RESEARCH AT MARINE STATIONS 80th anniversary of the Nikolai Pertsov White Sea Biological Station, MOSCOW, Russia
25. 26-30 May, 2019. The Crustacean Society Mid-Year Meeting, Hong Kong

List of main publications of Gregory A. Kolbasov
Total number of publications at present:
119 publications including 3 monographs and 7 chapters in joint monographs

1. Kolbasov, G.A., 1990a. Acasta pertusa sp.n. (Cirripedia,Thoracica) from the Red Sea. Zoologicheskiy zhurnal, 69(9): 142-145. (in Russian, with English abstract).
2. Kolbasov, G.A., 1990b. Acasta tabachniki (Cirripedia, Thoracica) from the South China Sea. Zoologicheskiy zhurnal, v.69(4): 135-137.(in Russian, with English abstract).
3. Kolbasov, G.A., 1991. New species of the genus Acasta (Cirripedia, Thoracica) from Western Pacific. Zoologicheskiy zhurnal, v.70(7):32-38.(in Russian, with English abstract).
4. Kolbasov, G.A., 1992a. Two new species of the genus Acasta (Cirripedia, Thoracica) from the South-western part of the Indian Ocean. Zoologicheskiy zhurnal, v.71(1):140-145. (in Russian, with English abstract).
5. Kolbasov, G.A., 1992b. New species of the genus Acasta (Cirripedia, Thoracica) from the South-western part of the Indian Ocean. Zoologicheskiy zhurnal, v.71(11):24-32. (in Russian, with English abstract).
6. Kolbasov, G.A., 1992c. Acasta spongiteformis (Cirripedia, Thoracica) from the South-west Indian Ocean. Zoologicheskiy zhurnal, v.71(12):133-136. (in Russian, with English abstract).
7. Kolbasov, G.A., 1993. Revision of the genus Acasta Leach (Cirripedia: Balanoidea). Zool. Jour. Linn. Soc., 109:395-427.
8. Kolbasov, G.A., 1996. The significance of symbiosis in the evolution of sessile barnacles (Cirripedia: Balanoidea). Arthropoda Selecta, 5 (1/2): 3-16.
9. Zevina, G.B., Kolbasov, G.A., 1997. Revision of the family Pagurolepadidae (Cirripedia, Lepadomorpha). Russian Journal of Zoology, 1(3): 281-291 (Translated from Zoologicheskii Zhurnal, 176(9): 992-1003. -an original Russian text).
10. Kolbasov, G.A., 1998. A new species of the genus Lithoglyptes (Cirripedia: Acrothoracica) from Indian Ocean (Isl. Java). Arthropoda Selecta, Vol.7 No. 3: 175-178.
11. Ilan, M., Loya, Y., Kolbasov, G.A., Brikner, I., 1999. Sponge inhabiting barnacles from coral reefs. Mar. Biol., 133: 709-716.
12. Kolbasov, G.A., 1999. The external mantle morphology of burrowing barnacles of the families Lithoglyptidae and Cryptophialidae (Cirripedia, Acrothoracica). Proceedings of the Fourth International Crustacean Congress, 1998, Amsterdam, the Netherlands, vol.I: 139-149.
13. Kolbasov, G.A., 1999. A new species of burrowing barnacles of the genus Lithoglyptes (Cirripedia, Acrothoracica, Lithoglyptidae) from the Maldives. Zoologicheskiy zhurnal, v.78(9): 1125-1128. (in Russian, with English abstract).
14. Kolbasov, G.A., Zevina, G.B., 1999. A new species of Paralepas (Cirripedia: Heteralepadidae) symbiotic with Xenophora (Mollusca: Gastropoda) with the first complemental male known for the family. Bull. Mar. Sci., 64(3):391-398.
15. Kolbasov, G.A., Høeg, J.T., Elfimov, A.S., 1999. Scanning electron microscopy of acrothoracican cypris larvae (Crustacea, Thecostraca, Cirripedia, Acrothoracica, Lithoglyptidae).Contrib. Zool., 68(3): 143-160.
16. Kolbasov, G.A., 2000. Lithoglyptes cornutus, new species (Cirripedia: Acrothoracica), a boring barnacle from Seychelles, with some data on the ultrastructure. Hydrobiologia, 438(1-3): 185-191.
17. Kolbasov, G.A., 2000 A new species of the genus Lithoglyptes (Cirripedia, Acrothoracica, Lithoglyptidae) offshore island Nauru (Micronesia). Zoologicheskiy zhurnal, v.79(3): 372-376. (in Russian, with English abstract).
18. Kolbasov, G.A., 2000. External morphology of the burrowing barnacle Lithoglyptes bicornis (Crustacea:Cirripedia: Acrothoracica: Lithoglyptidae). Hydrobiologia, 438(1-3): 193-204.
19. Kolbasov, G.A., 2000. New species, Cryptophialus hoegi (Cirripedia, Acrothoracica, Cryptophialidae) from the Aden Gulf. Description of female and dwarf male ultrastructure. Zoologicheskiy zhurnal, v.79(9):1027-1035. (in Russian, with English abstract).
20. Kolbasov, G.A., 2000. Calcareous plates in barnacles of the superorder Acrothoracica, morphology and functions, homology with calcareous plates of barnacles of the superorder Thoracica (Thecostraca, Cirripedia). Zoologicheskiy zhurnal, v.79(11): 1284-1289. (in Russian, with English abstract).
21. Kolbasov, G.A., 2000. A new species of the genus Lithoglyptes (Crustacea: Cirripedia: Acrothoracica) from the Fiji Sea, with some data on its ultrastructure. Arthropoda Selecta, 9(2): 85-89.
22. Kolbasov, G.A., Høeg, J.T., 2000. External morphology of females in the burrowing barnacles Lithoglyptes mitis and L. habei (Lithoglyptidae) and the phylogenetic position of the Cirripedia Acrothoracica (Crustacea: Thecostraca). Arthropoda Selecta, 9(1): 13-27.
23. Zevina, G.B., Kolbasov, G.A., 2000. Barnacles of the genus Heteralepas (Thecostraca, Cirripedia, Thoracica) from the Canary Islands and the Azores. Description of mantle ultrastructure. Zoologicheskiy zhurnal, v.79 (11): 1275-1283. (in Russian, with English abstract).
24. Kolbasov, G.A., 2001. A new species of the burrowing barnacle Weltneria bekae (Cirripedia, Acrothoracica) from Papua (New Guinea). Description of external morphology. Zoologicheskiy zhurnal, v.80(10): 1-10. (in Russian, with English abstract).
25. Kolbasov, G.A., Høeg, J.T., 2001. Exterior morphology of the burrowing barnacle Trypetesa lampas (Cirripedia, Acrothoracica). Zoologicheskiy zhurnal, v.80(5): 525-533. (in Russian, with English abstract).
26. Kolbasov, G.A., Høeg, J.T., 2001. External morphology of cypris larvae of two species of Trypetesa Norman, 1903 (Crustacea, Thecostraca, Cirripedia, Acrothoracica, Trypetesidae). Arthropoda Selecta, 10(2): 87-92.
27. Kolbasov, G.A., Pegova, A.N., 2001. Chromosome set in crustaceans of the subclass Facetotecta (Crustacea, Thecostraca) from the White Sea. Zoologicheskiy zhurnal, v.80(12): 1444-1451. (in Russian, with English abstract).
28. Kolbasov, G.A., 2002. Cuticular structures of some acrothoracican dwarf males (Crustacea: Thecostraca: Cirripedia: Acrothoracica). Zool. Anz., 241: 85-94.
29. Kolbasov, G.A., 2002. A new species of burrowing barnacles, Kochlorine grebelnii, from the Cape Verde Islands. Description of external ultrastructure of the genus Kochlorine (Thecostraca, Cirripedia, Acrothoracica). Zoologicheskiy zhurnal, v.81(5): 529-546. (in Russian, with English abstract).
30. Høeg, J.T., Kolbasov, G.A., 2002. Lattice organs in y cyprids of the Facetotecta and their significance in the phylogeny of the Crustacea Thecostraca. Acta Zoologica, 83: 67-79 .
31. Pérez-Losada, M., Høeg, J.T., Kolbasov, G.A., Crandall, K.A., 2002. Reanalysis of the relationships among the Cirripedia and the Ascothoracida and the phylogenetic position of the Facetotecta (Maxillopoda: Thecostraca) using 18S rDNA sequences. J. Crustacean Biol., 22(3): 661-669.
32. Kolbasov, G.A., Høeg, J.T., 2003. Facetotectan larvae from the White Sea with description of a new species (Crustacea: Thecostraca). Sarsia, 88: 1-15.
33. Rybakov, A.V., Høeg, J.T., Jensen, P.G., Kolbasov, G.A., 2003. The chemoreceptive lattice organs in cypris larvae develop from naupliar setae (Thecostraca: Cirripedia, Ascothoracida and Facetotecta). Zoologischer Anzeiger, 242: 1-20.
34. Kolbasov, G.A., 2004 A new burrowing barnacle species and external ultrastructure in adults of the family Cryptophialidae (Thecostraca, Cirripedia, Acrothoracica). Zoologicheskiy zhurnal, v.83(1): 16-32. (in Russian, with English abstract).
35. Kolbasov, G.A., 2004. A new species of crustaceans of the genus Gorgonolaureus (Crustacea, Thecostraca, Ascothoracida) – parasites of gorgonians from Vietnam. Zoologicheskiy zhurnal, v.83(5): 536-548. (in Russian, with English abstract).
36. Kolbasov, G.A., Newman, W.A., 2005. Revision of the Lithoglyptidae sensu Tomlinson, 1969 and Lithoglyptes Aurivillius, 1892 (Cirripedia, Acrothoracica), including a new species from Bermuda. Zootaxa, 1013: 35-64.
37. Kolbasov, G.A, Buckeridge, J.S, Høeg, J.T, Newman, W.A. 2005 Galina Benizianovna Zevina - February 12, 1926-September 24, 2002. Journal of Crustacean Biology, 25 (2): 309-314.
38. Kolbasov, G.A., 2007. External morphology of parasitic crustaceans of genus Dendrogaster (Thecostraca, Ascothoracida). Zoologicheskiy Zhurnal, 86(6): 654-670. (in Russian with English abstract).
39. Kolbasov, G.A., J.T. Høeg. 2007. Cypris larvae of the acrothoracican barnacles (Thecostraca, Cirripedia, Acrothoracica). Zoologischer Anzeiger, 246(2): 127-151.
40. Kolbasov, G.A., 2007. Microscopic parasitic crustaceans of class Tantulocarida. Morphology, systematics and development. Uchenye zapiski of Kazan State University, Ser. Nat. Sciences, V.149, B. 3: 89-92.
41. Kolbasov, G.A., M.J. Grygier, V.N. Ivanenko, A.A. Vagelli, 2007. A new species of the y-larva genus Hansenocaris Itô, 1985 (Crustacea: Thecostraca: Facetotecta) from Indonesia, with a review of y-ciprids and a key to all their described species. The Raffles Bulletin of Zoology, 55(2): 343-353.
42. Kolbasov, G.A., M.J. Grygier, J.T. Høeg, W. Klepal 2008. External morphology of ascothoracid-larvae of the genus Dendrogaster (Crustacea, Thecostraca, Ascothoracida), with remarks on the ontogeny of the lattice organs. Zoologischer Anzeiger, 247: 159-183.
43. Kolbasov, G.A., Sinev, A.Yu., Tchesunov, A.V., 2008. External morphology of Arcticotantulus pertzovi – microscopic crustacean parasite from the White Sea (Crustacea, Tantulocarida). Zoologicheskiy zhurnal, 87 (12): 1437-1452. (in Russian with English abstract)
44. Kolbasov G.A., 2009. Subclass Cirripedia (superorder Thoracica). In Illustrated keys to free-living invertebrates of Eurasian Arctic seas and adjacent deep waters, Vol. 1, eds. S.V. Vasilenko and V.V. Petryashov, Alaska, USA: 49-60
45. Kolbasov, G.A., 2009. Acrothoracica, burrowing crustaceans. Moscow, KMK press, 452 pp.
46. Savchenko A.S., Kolbasov G.A., 2009. Serratotantulus chertoprudae gen. et sp. n. (Crustacea, Tantulocarida, Basipodellidae) –a new tantulocaridan from the abyssal depths of the Indian Ocean. Integrative and Comparative Biology, Symposium ‘‘The Biology of the Parasitic Crustacea’’, January 3–7, 2009, Boston, Massachusetts, 1-8.
47. Høeg J.T., Pérez-Losada M., Glenner H., Kolbasov G.A., Crandall K.A., 2009. Evolution of morphology, ontogeny and life cycles within the Crustacea Thecostraca. Arthropod Systematics and Phylogeny, 67(2): 199-217.
48. Kolbasov G.A., Savchenko A.S. 2010. Microdajus tchesunovi sp. n. (Tantulocarida, Microdajidae) – A new crustacean parasite from the White Sea. Experimental Parasitology. Vol. 125: 13-22.
49. Chan B.K.K, Cheang C.C., Chen I-H., Kolbasov G.A. 2013. Acrothoracican barnacles (Lithoglyptida) in Taiwan, including the taxonomic status of Balanodytes taiwanus Utinomi, 1950 and cryptic diversity of Auritoglyptes bicornis (Aurivillius, 1892). Zootaxa, 3694 (3): 221–239.
50. Benny K.K. Chan, Gregory A. Kolbasov, Chi-Chiu Cheang. 2012. Cryptic diversity of the acrothoracican barnacle Armatoglyptes taiwanus in the Indo-Pacific waters, with description of a new species from the Mozambique Channel collected from the French MAINBAZA Cruise. Zoosystema, 34(1): 5-20.
51. Petrunina A.S., Kolbasov G.A. 2012. Morphology and ultrastructure of definitive males of Arcticotantulus pertzovi and Microdajus tchesunovi (Crustacea; Tantulocarida). Zoologischer Anzeiger, 251: 223-236.
52. Petrunina A.S., Neretina T.V., Mugue N.S., Kolbasov G.A. 2013. Tantulocarida versus Thecostraca: inside or outside? First attempts to resolve phylogenetic position of Tantulocarida using gene sequences. Journal of Zoological Systematics and Evolutionary Research. 1-9, doi: 10.1111/jzs.12045
53. Kolbasov G.A., Elfimov A.S., Høeg J.T. 2013. External morphology of barnacle cypris larvae in the family Poecilasmatidae (Cirripedia: Thoracica: Pedunculata): Toward a template for scoring cypris characters. Zoologischer Anzeiger, 252: 522-535.
54. Chan B.K.K., Kolbasov G., Hirose M., Mezaki T., Suwa R. 2014. Biodiversity and biogeography of the coral boring barnacles of the genus Berndtia (Cirripedia: Acrothoracica) in the West Pacific, with description of three new species. Journal of Natural History, Vol. 48, Nos. 25–26, 1503–1541,
55. Chan B.K.K., Hsieh W.-P., Kolbasov G.A. 2014. Crustacean Fauna of Taiwan: Barnacles, Volume III Cirripedia: Acrothoracica. Publ. Biodiversity Research Centre, Academia Sinica, Taipei, Taiwan, 107 pp.
56. Høeg J.T., Chan B.K.K., Kolbasov G.A., Grygier M.J. 2014. Facetotecta. In: Martin, J., Olesen, J., and Høeg, J.T. (Eds) Atlas of Crustacean Larvae. Johns Hopkins University Press, 100-103. ISBN-13: 978-1421411972
57. Høeg J.T., Chan B.K.K., Kolbasov G.A., Grygier M.J. 2014. Ascothoracida. In: Martin, J., Olesen, J., and Høeg, J.T. (Eds) Atlas of Crustacean Larvae. Johns Hopkins University Press, 104-106. ISBN-13: 978-1421411972
58. Kolbasov G.A., Chan B.K.K., Høeg J.T. 2014. Acrothoracica. In: Martin, J., Olesen, J., and Høeg, J.T. (Eds) Atlas of Crustacean Larvae. Johns Hopkins University Press, 107-110. ISBN-13: 978-1421411972
59. Kolbasov G.A., Chan B.K.K., Petrunina A.S. 2015. Gorgonolaureus bicornutus sp. nov (Crustacea: Thecostraca: Ascothoracida) from off South-East Taiwan with notes on morphology and distribution. Zootaxa, 3972 (3): 328–342.
60. Kolbasov G.A. 2015. Superorder Acrothoracica. Revista IDE@ - SEA, n 101B (30-06-2015): 1–6. ISSN 2386-7183, Ibero Diversidad Entomológica @ccesible,
61. Kolbasov G.A. 2015. Infraclass Ascothoracida. Revista IDE@ - SEA, n 100B (30-06-2015): 1–6. ISSN 2386-7183, Ibero Diversidad Entomológica @ccesible,
62. Lin H.Ch., Kolbasov G., Chan B.K.K. 2016. Phylogenetic relationships of Darwin's "Mr Arthrobalanus": the burrowing barnacles (Cirripedia: Acrothoracica). Molecular Phylogenetics and Evolution, 100 (2016): 292-302.
63. Meng-Chen Yu, Gregory A. Kolbasov, Benny K.K. Chan. 2016. New species of sponge inhabiting barnacle of genus of Bryozobia (Archaeobalanidae, Bryozobiinae) in the West Pacific. ZooKeys, 571: 1–20. doi: 10.3897/zookeys.571.6894.
64. Kolbasov Gregory A., Achituv Yair Chan Benny K.K. Molodtsova T.N. 2016. Revision of the coral-inhabiting genus Conopea (Cirripedia: Archaeobalanidae) with description of two new species of the genera Conopea and Acasta. Zootaxa, 4178 (2): 182–208.
65. Benny K. K. Chan, Gregory A. Kolbasov, and Jens T. Høeg. 2016. Collecting and processing thoracican, acrothoracican and rhizocephalan Cirripedes. Journal of Crustacean Biology, 36(5): 752-760. DOI: 10.1163/1937240X-00002469.
66. Meng-Chen Yu, Gregory A. Kolbasov, Andrew M. Hosie, Tse-Min Lee & Benny K.K. Chan. 2017. Descriptions of four new sponge-inhabiting barnacles (Thoracica: Archaeobalanidae: Acastinae). Zootaxa 4277 (2): 151–198 (10.11646/zootaxa.4277.2.1).
67. Gregory A. Kolbasov, Benny K.K. Chan & Yu-Rong Cheng. 2017. Weltneria acanthostoma sp. nov., a burrowing barnacle (Cirripedia: Acrothoracica) from the deep-waters of the South China Sea. Zootaxa, 4290 (3): 591-599 (10.11646/zootaxa.4290.3.12)
68. Meng-Chen Yu, Chan Benny K.K, Yair Achituv, Kolbasov Gregory A., 2017. Four new sponge-inhabiting barnacles of the genus Acasta (Thoracica: Archaeobalanidae: Acastinae) from the Indo-Pacific. Raffles Bulletin of Zoology, 65, 585-615.
69. Kolbasov G.A., Newman W.A. 2018. A new species of Synagoga (Crustacea: Thecostraca: Ascothoracida) parasitic on an antipatharian from the Azores and Cape Verde Islands, with notes on its morphology, sexuality, host specificity, and biogeography. Marine Biodiversity, 49(3): 1101–1122.
70. Petrunina A.S., Høeg J.T., Kolbasov G.A. 2018. Anatomy of the Tantulocarida: first results obtained using TEM and CLSM. Part I: tantulus larva. Organisms Diversity and Evolution, 18 (4): 459-477 (10.1007/s13127-018-0376-4)
71. Kolbasov G.A., Petrunina A.S. 2018. The family Ascothoracidae Grygier, 1987, a review with descriptions of new abyssal taxa parasitizing ophiuroids and remarks on the invalidity of the genus Parascothorax Wagin, 1964 (Crustacea: Thecostraca: Ascothoracida). Marine Biodiversity, 1-31(10.1007/s12526-018-0921-6)
72. Meng-Chen Yu, Gregory A. Kolbasov, Jens T. Høeg, Benny K.K. Chan. 2019. Crustacean-sponge symbiosis: collecting and maintaining sponge-inhabiting barnacles (Cirripedia: Thoracica: Acastinae) for studies on host specificity and larval biology. Journal of Crustacean Biology (2018) 1–11. doi:10.1093/jcbiol/ruz025
73. Kolbasov Gregory A., Petrunina Alexandra S., Ming-Jay Ho, Chan Benny K.K. 2019. A new species of Synagoga (Crustacea, Thecostraca, Ascothoracida) parasitic in an antipatharian from Green Island, Taiwan, with notes on its morphology. ZooKeys, 876: 55-85. DOI: 10.3897/zookeys.876.35443
74. Alexandra S. Petrunina, Gregory A. Kolbasov, Pedro Martinez Arbizu. 2019. Anatomy of the free tantulus larva (Crustacea: Tantulocarida) studied with confocal laser scanning microscopy: An extreme case of miniatuarisation in the Arthropoda. Progress in Oceanography, Volume 178, November 2019, 102190.
75. Gregory A. Kolbasov, Alexandra S. Petrunina, Jørgen Olesen, Ming-Jay Ho, Benny K. K. Chan, Mark J. Grygier. 2020. A new species of Sessilogoga parasitic in an antipatharian from Green Island, Taiwan, with notes on its nauplius larvae and the synapomorphies and apparent gonochorism of the genus (Crustacea: Thecostraca: Ascothoracida). Marine Biodiversity, 50, 43.
76. Yu M.-C., Dreyer N., Kolbasov G.A., Høeg J.T., Chan B.K.K. 2020. Sponge symbiosis is facilitated by adaptive evolution of larval sensory and attachment structures in barnacles. Proceedings of Royal Society B. 28720200300
77. Meng-Chen Yu, Takefumi Yorisue, Gregory A. Kolbasov, Benny K.K. Chan. 2020. A new species of sponge-inhabiting barnacle Neoacasta (Archaeobalanidae, Acastinae) in Kyushu, Japan. Crustaceana, 93(11-12): 1515–1532. DOI:
78. Meng-Chen Yu, Benny K K Chan, Gregory A Kolbasov, Monthon Ganmanee. 2020. Biodiversity and host specificity of sponge-associated barnacles (Cirripedia: Thoracica) in Thailand, Journal of Crustacean Biology, ruaa080,
79. Chan B.K.K., Dreyer N., Gale A.S., Glenner H., Ewers-Saucedo C., Pérez-Losada M., Kolbasov G.A., Crandall K.A., Høeg J.T. The evolutionary diversity of barnacles, with an updated classification of fossil and living forms Zoological Journal of the Linnean Society, 2021, XX, 1–58. DOI: 10.1093/zoolinnean/zlaa160
80. Kolbasov, G.A., Petrunina, A.S., Dreyer, N. Prudkovky A.A., Olesen J., Chan B.K.K., Høeg J.T. 2021. Naupliar development of Facetotecta (Crustacea: Thecostraca) and the nature of the first nauplius instar in the Crustacea - pro et contra the Hexanauplia concept. Org. Divers. Evol. 21, 209–230 (2021).


June 2019
Summer course on embryology of marine invertebrates


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